Henderson (2010) estimated the body masses of 14 species of pterosaur using three-dimensional digital models. The largest and most troubling pterosaur
was Quetzalcoatlus northropi. The estimated mass of 544 kg was considered “astonishingly high”. However, it was noted that the previous low mass estimates of 64 kg and 70 kg required “more than 90% of the body filled with air” so were difficult to accept biologically. Henderson (2010, p783) suggested that Quetzalcoatlus northropi might have been flightless since,
“Accepting this possibility releases us from the requirement to generate, and accept, unrealistically low masses and body densities for such large animal. It also frees us from the mental gymnastics required to generate an anatomy with sufficient muscle mass and power to be able to fly
when possibly weighing more than thirty times that of the heaviest, living birds such as the 16-kg
Kori Bustard (Ardeotis kori) and the Great Bustard (Otis tarda), which may attain 22 kg in some cases. These birds seem to be at the upper mass limit for flying given their apparent difficulty in taking off (Alexander, 1998). A study of the scaling relationships between body mass and the
flapping and gliding flight styles in procellariform birds—albatrosses and their closest relatives—predicts an upper body mass limit of 41 kg for this type of flying (Sato et al., 2009).”

亨德森（2010）利用三维数字模型估算了14种翼龙的体重。其中体型最大、也最令人困惑的是风神翼龙（Quetzalcoatlus northropi），其估算体重达544千克，被认为“高得惊人”。但他指出，此前64千克和70千克的低体重估算“需要身体90%以上的空间充满空气”，从生物学角度来看难以成立。因此，亨德森（2010，p783）认为风神翼龙可能是不会飞的，因为：
“接受这种可能性，我们就无需再去推测并接受这种大型动物不切实际的低体重和身体密度。这也让我们不必再绞尽脑汁地去构想一种具备足够肌肉量和动力、却可能比现存最重的鸟类（如16千克的灰颈鸨和某些情况下可达22千克的大鸨）重三十倍以上的解剖结构，以便解释它能够飞行的原因。考虑到这些鸟类起飞时明显存在困难，它们似乎已处于飞行体重的上限（亚历山大，1998）。一项关于鹱形目鸟类（信天翁及其近亲）体重与扑翼飞行、滑翔飞行方式之间比例关系的研究预测，此类飞行方式的体重上限为41千克（佐藤等人，2009）。”

省流，Henderson (2010)认为风神体重有544kg，但他认为这个体重的风神飞不起来

所以他认为风神是纯地栖的吗

为什么飞行动物的体重上限这么低，是跟能量消耗还有什么有关呢

翼龙不会飞跟棘龙会飞的可能性哪个大

本应如此

不可靠……要是真完全不会飞，那形态上不可能还保留那么多使应飞行的特征，不光是翼膜

不可能滑翔吧？